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The Cytoskeleton course addresses advanced topics in the mechanochemical systems that govern and drive cell shape, tissue morphogenesis and integrity, mechanosensing, and the integration chemical signaling and mechanical inputs (force, stress, adhesion). These cellular systems are at the crux of healthy development and disease. We take a no-holds-barred approach in order to have advanced, in depth discussions of the topic. We will embrace molecular, biochemical, biophysical and computational logic and approaches in our thinking during the course.